Dr Thomas Woolley

2026

• Farlie, F. et al., 2026. Balancing science and sustainability in ART: An international survey of green issues in IVF laboratories. Reproductive BioMedicine Online 53 (4) 105804. (10.1016/j.rbmo.2026.105804)
• Woolley, T. 2026. Turing bifurcations on elliptical domains: A deviation from rectangular and circular limits. International Journal of Applied and Computational Mathematics 12 (2) 25. (10.1007/s40819-026-02100-2)
• Bansal, N. , Kaouri, A. and Woolley, T. 2026. Reducing size bias in epidemic network modelling. Journal of Theoretical Biology 618 112314. (10.1016/j.jtbi.2025.112314)
• Woolley, T. 2026. Mind the gap: A solution to the robustness problem of Turing patterns through patterning mode isolation. AppliedMath 6 (1) 3.
• Woolley, T. and Klika, V. 2026. Role of dimension and geometry: non-uniqueness in the pseudo-one-dimensional limit for Turing patterns on parallelograms. European Journal of Applied Mathematics

2025

• Jabi, W. et al. 2025. 3D topological modeling and multi-agent movement simulation for viral infection risk analysis. Architectural Science Review (10.1080/00038628.2025.2603534)
• Woolley, T. 2025. Breaking free: motivating mathematics through escape rooms. MSOR Connections 24 (1), pp.75-83.
• Lewis, T. et al., 2025. Searching for model structures on murine neonatal sepsis data using genetic programming. Presented at: GECCO 2025 Málaga, Spain 14-18 April 2025. Published in: Ochoa, G. and Filipič, B. eds. GECCO '25 Companion: Proceedings of the Genetic and Evolutionary Computation Conference Companion. ACM. , pp.863-866. (10.1145/3712255.3726680)
• Noone, C. et al., 2025. Critically appraising the Cass Report: Methodological flaws and unsupported claims. BMC Medical Research Methodology 25 128. (10.1186/s12874-025-02581-7)
• Woolley, T. 2025. Bespoke Turing patterns with specific nonlinear properties. Proceedings of the Royal Society A: Mathematical, Physical and Engineering Sciences 481 (2312) 20250030. (10.1098/rspa.2025.0030)
• Hakami, A. et al. 2025. Graft ischemia post cell transplantation to the brain: Glucose deprivation as the primary driver of rapid cell death. Neurotherapeutics 22 (2) e00518. (10.1016/j.neurot.2024.e00518)
• Palmer, G. et al., 2025. Use and understanding of AI in the ART laboratory: an international survey. Reproductive BioMedicine Online 50 (3) 104435. (10.1016/j.rbmo.2024.104435)
• Woolley, T. 2025. Pattern formation on regular polygons and circles. Journal of Nonlinear Science 35 5. (10.1007/s00332-024-10096-6)
• Mahashri, N. , Woolley, T. E. and Muthusamy, C. 2025. Linear coupling of patterning systems can have nonlinear effects. Physical Review E 111 (1) 014224. (10.1103/physreve.111.014224)

2024

• Xue, Y. et al. 2024. Modelling indoor airborne transmission combining architectural design and people movement using the VIRIS simulator and web app. Scientific Reports 14 28220. (10.1038/s41598-024-79525-6)
• Tseng, C. et al., 2024. Gap junctions in Turing-type periodic feather pattern formation. PLoS Biology 22 (5) e3002636. (10.1371/journal.pbio.3002636)
• Henley, L. et al., 2024. A simple and fast method for estimating bat roost locations. Royal Society Open Science 11 (4) 231999. (10.1098/rsos.231999)
• Gallagher, K. , Ostler, T. and Woolley, T. 2024. Retinal oxygenation with conventional 100ms vs short-pulse pan-retinal laser photocoagulation. Ophthalmic Surgery, Lasers and Imaging Retina 55 (1), pp.40-45. (10.3928/23258160-20231114-01)
• Henley, L. et al., 2024. Bat motion can be described by leap frogging. Bulletin of Mathematical Biology 86 16. (10.1007/s11538-023-01233-5)

2023

• Moore, J. W. , Dale, T. C. and Woolley, T. E. 2023. Modelling polarity-driven laminar patterns in bilayer tissues with mixed signalling mechanisms. SIAM Journal on Applied Dynamical Systems 22 (4), pp.2945-2990. (10.1137/22M1522565)
• Smith, C. J. et al., 2023. Unravelling the clinical co-morbidity and risk factors associated with agenesis of the corpus callosum. Journal of Clinical Medicine 12 (11) 3623. (10.3390/jcm12113623)
• Oruganti, S. et al., 2023. Immune and metabolic markers for identifying and investigating severe Coronavirus disease and Sepsis in children and young people (pSeP/COVID ChYP study): protocol for a prospective cohort study. BMJ Open 13 e067002. (10.1136/bmjopen-2022-067002)
• Glover, J. D. et al., 2023. The developmental basis of fingerprint pattern formation and variation. Cell 186 , pp.1-17. (10.1016/j.cell.2023.01.015)

2022

• Moore, S. C. , Woolley, T. E. and White, J. 2022. An exploration of the multiplicative effect of "Other people" and other environmental effects on violence in the night-time environment. International Journal of Environmental Research and Public Health 19 (24) 16963. (10.3390/ijerph192416963)
• May, S. et al. 2022. Modification of diet to reduce the stemness and tumourigenicity of murine and human intestinal cells. Molecular Nutrition & Food Research 66 (19) 2200234. (10.1002/mnfr.202200234)
• Ghazal, P. et al. 2022. Corrigendum to "Challenging molecular dogmas in human sepsis using mathematical reasoning" [EBioMedicine 80 (2022) 104031]. EBioMedicine 85 104331. (10.1016/j.ebiom.2022.104331)
• Moore, J. W. , Dale, T. C. and Woolley, T. E. 2022. Polarity driven laminar pattern formation by lateral-inhibition in 2D and 3D bilayer geometries. IMA Journal of Applied Mathematics 87 (4), pp.568-606.
• Woolley, T. 2022. Boundary conditions cause different generic bifurcation structures in Turing systems. Bulletin of Mathematical Biology 84 (9) 101.
• Ghazal, P. et al. 2022. Challenging molecular dogmas in human sepsis using mathematical reasoning. EBioMedicine 80 104031. (10.1016/j.ebiom.2022.104031)
• Woolley, T. , Hill, W. and Hogan, C. 2022. Accounting for dimensional differences in stochastic domain invasion with applications to precancerous cell removal. Journal of Theoretical Biology 541 111024. (10.1016/j.jtbi.2022.111024)
• Robinson, J. et al., 2022. The association of neurodevelopmental abnormalities, congenital heart and renal defects in a Tuberous Sclerosis Complex patient cohort. BMC Medicine 20 123. (10.1186/s12916-022-02325-0)
• Buze, M. , Woolley, T. E. and Mihai, L. A. 2022. A stochastic framework for atomistic fracture. SIAM Journal on Applied Mathematics 82 (2), pp.526-548. (10.1137/21M1416436)

2021

• Kasemeier-Kulesa, J. C. et al., 2021. The embryonic trunk neural crest microenvironment regulates the plasticity and invasion of human neuroblastoma via TrkB signaling. Developmental Biology 480 , pp.78-90. (10.1016/j.ydbio.2021.08.007)
• Moore, J. W. et al. 2021. A general computational framework for COVID-19 modelling with applications to testing varied interventions in education environments. COVID 1 (4), pp.674-703. (10.3390/covid1040055)
• Ostler, T. et al. 2021. Vitrifying multiple embryos in different arrangements does not alter the cooling rate. Cryobiology 103 , pp.22-31. (10.1016/j.cryobiol.2021.10.001)
• Suarez-Berumen, K. et al., 2021. Pannexin 1 regulates skeletal muscle regeneration by promoting bleb-based myoblast migration and fusion through a novel lipid based signaling mechanism. Frontiers in Cell and Developmental Biology 9 736813. (10.3389/fcell.2021.736813)
• Simonović, J. and Woolley, T. 2021. Generalised S-System-Type Equation: Sensitivity of the deterministic and stochastic models for bone mechanotransduction. Mathematics 9 (19) 2422. (10.3390/math9192422)
• Hill, W. et al. 2021. EPHA2-dependent outcompetition of KRASG12D mutant cells by wild-type neigbors in the adult pancreas. Current Biology 31 (12), pp.2550-2560. E5. (10.1016/j.cub.2021.03.094)
• Woolley, T. E. , Krause, A. L. and Gaffney, E. A. 2021. Bespoke Turing systems. Bulletin of Mathematical Biology 83 41. (10.1007/s11538-021-00870-y)
• Harn, H. I. et al., 2021. Symmetry breaking of tissue mechanics in wound induced hair follicle regeneration of laboratory and spiny mice. Nature Communications 12 2595. (10.1038/s41467-021-22822-9)
• Harper, P. , Moore, J. and Woolley, T. 2021. Covid-19 transmission modelling of students returning home from university. Health Systems 10 (1), pp.31-40. (10.1080/20476965.2020.1857214)
• Moore, J. et al. 2021. Further development of spinal cord retreatment dose estimation: including radiotherapy with protons and light ions. International Journal of Radiation Biology 97 (12), pp.1657-1666. (10.1080/09553002.2021.1981554)

2020

• Krause, A. et al., 2020. Turing patterning in stratified domains. Bulletin of Mathematical Biology 82 136. (10.1007/s11538-020-00809-9)
• Henley, L. et al. 2020. Long term environmental implications of social distancing on public transport emissions. Project Report.[Online].London: UK Government. Available at: https://committees.parliament.uk/writtenevidence/8995/pdf/.
• Mihai, L. A. , Woolley, T. and Goriely, A. 2020. Likely cavitation and radial motion of stochastic elastic spheres. Nonlinearity 33 (5) 1987. (10.1088/1361-6544/ab7104)
• Krause, A. et al., 2020. From one pattern into another: analysis of Turing patterns in heterogeneous domains via WKBJ. Interface 17 (162)(10.1098/rsif.2019.0621)
• Adamer, M. F. et al., 2020. Coloured noise from stochastic inflows in reaction-diffusion systems. Bulletin of Mathematical Biology 82 44. (10.1007/s11538-020-00719-w)
• Harper, P. , Moore, J. and Woolley, T. 2020. Secondary household Covid-19 transmission modelling of students returning home from university. n/a
• Viglialoro, G. and Woolley, T. E. 2020. Solvability of a Keller-Segel system with signal-dependent sensitivity and essentially sublinear production. Applicable Analysis 99 (14), pp.2507-2525. (10.1080/00036811.2019.1569227)

2019

• Fitt, D. et al. 2019. Uncertainty quantification of elastic material responses: testing, stochastic calibration and Bayesian model selection. Mechanics of Soft Materials 1 13. (10.1007/s42558-019-0013-1)
• Budia, I. et al., 2019. Radiation protraction schedules for low-grade gliomas: A comparison between different mathematical models. Interface 16 (161) 20190665. (10.1098/rsif.2019.0665)
• Picco, N. and Woolley, T. 2019. Time to change your mind? Modelling transient properties of cortex formation highlights the importance of evolving cell. Journal of Theoretical Biology 481 , pp.110-118. (10.1016/j.jtbi.2018.08.019)
• Maini, P. K. and Woolley, T. 2019. The Turing model for biological pattern formation. In: Bianchi, A. et al., The Dynamics of Biological Systems. Mathematics of Planet Earth Springer. , pp.189-204.
• Mihai, L. A. et al. 2019. Likely cavitation in stochastic elasticity. Journal of Elasticity 137 (1), pp.27-42. (10.1007/s10659-018-9706-1)
• Picco, N. et al., 2019. A mathematical insight into cell labelling experiments for clonal analysis. Journal of Anatomy 235 (3), pp.687-696. (10.1111/joa.13001)
• Mihai, L. A. , Woolley, T. and Goriely, A. 2019. Likely chirality of stochastic anisotropic hyperelastic tubes. International Journal of Non-Linear Mechanics 114 , pp.9-20. (10.1016/j.ijnonlinmec.2019.04.004)
• Mihai, L. A. et al. 2019. Likely equilibria of stochastic hyperelastic spherical shells and tubes. Mathematics and Mechanics of Solids 24 (7), pp.2066-2082. (10.1177/1081286518811881)
• Simonovic, J. and Woolley, T. 2019. Deterministic and stochastic parameter analysis of the bone cell population model. Presented at: 8th International Conference on Computational Bioengineering (ICCB2019) Belgrade, Serbia 4-6 September 2019.
• Simonovic, J. and Woolley, T. 2019. Equation of osteocyte activity in the bone cell population model. Presented at: ACTC Advances in Cell and Tissue Cultures, Cardiff University School of Biosciences Cardiff, UK 4-5th June 2019.
• Mihai, L. A. , Woolley, T. E. and Goriely, A. 2019. Likely equilibria of the stochastic Rivlin cube. Philosophical Transactions of the Royal Society A: Mathematical, Physical and Engineering Sciences 377 (2144) 20180068. (10.1098/rsta.2018.0068)
• Weber, E. et al., 2019. Self-organizing hair peg-like structures from dissociated skin progenitor cells: New insights for human hair follicle organoid engineering and Turing patterning in an asymmetric morphogenetic field. Experimental Dermatology 28 (4), pp.355-366. (10.1111/exd.13891)
• Mihai, L. et al. 2019. Likely oscillatory motions of stochastic hyperelastic solids. Transactions of Mathematics and Its Applications 3 (1) tnz003. (10.1093/imatrm/tnz003)
• Simonovic, J. and Woolley, T. 2019. Study of bone cell population models of s-system type. Presented at: Organ-on-a-Chip Technologies Network – Learning and Collaborative Event on 8 April 2019 at QMUL London, UK 8-9 April 2019.
• Ho, W. K. W. et al., 2019. Feather arrays are patterned by interacting signalling and cell density waves. PLoS Biology 17 (2) e3000132. (10.1371/journal.pbio.3000132)
• Navajas Acedo, J. et al., 2019. Parallel control of mechanosensory hair cell orientation by the PCP and Wnt pathways. Nature Communications 10 3993. (10.1038/s41467-019-12005-y)

2018

• Viglialoro, G. and Woolley, T. E. 2018. Eventual smoothness and asymptotic behaviour of solutions to a chemotaxis system perturbed by a logistic growth. Discrete and Continuous Dynamical Systems - Series B 23 (8), pp.3023-3045. (10.3934/dcdsb.2017199)
• Kasemeier-Kulesa, J. C. et al., 2018. Predicting neuroblastoma using developmental signals and a logic-based model. Biophysical Chemistry 238 , pp.30-38. (10.1016/j.bpc.2018.04.004)
• Picco, N. et al., 2018. Mathematical modeling of cortical neurogenesis reveals that the founder population does not necessarily scale with neurogenic output. Cerebral Cortex 28 (7), pp.2540-2550. (10.1093/cercor/bhy068)
• Krause, A. L. et al., 2018. Heterogeneity induces spatiotemporal oscillations in reaction-diffusions systems. Physical Review E 97 052206. (10.1103/PhysRevE.97.052206)
• Woolley, T. et al. 2018. Changes in the retreatment radiation tolerance of the spinal cord with time after the initial treatment. International Journal of Radiation Biology 94 , pp.515-531. (10.1080/09553002.2018.1430911)
• Sanders, J. et al. 2018. PLCz induced Ca2+ oscillations in mouse eggs involve a positive feedback cycle of Ca2+ induced InsP3 formation from cytoplasmic PIP2. Frontiers in Cell and Developmental Biology 6 36. (10.3389/fcell.2018.00036)
• Viglialoro, G. and Woolley, T. 2018. Boundedness in a parabolic-elliptic chemotaxis system with nonlinear diffusion and sensitivity, and logistic source. Mathematical Methods in the Applied Sciences 41 (5), pp.1809-1824. (10.1002/mma.4707)
• Mihai, L. A. , Woolley, T. and Goriely, A. 2018. Stochastic isotropic hyperelastic materials: constitutive calibration and model selection. Proceedings of the Royal Society A 474 (2211) 201708. (10.1098/rspa.2017.0858)

2017

• Adamer, M. , Woolley, T. and Harrington, H. 2017. Graph-facilitated resonant mode counting in stochastic interaction networks. Journal of the Royal Society Interface 14 (137) 20170447. (10.1098/rsif.2017.0447)
• Woolley, T. E. 2017. Pattern production through a chiral chasing mechanism. Physical Review E 96 032401. (10.1103/PhysRevE.96.032401)
• Woolley, T. E. , Gaffney, E. A. and Goriely, A. 2017. Random blebbing motion: a simple model linking cell structural properties to migration characteristics. Physical Review E 96 012409. (10.1103/PhysRevE.96.012409)

• Hamilton, F. et al. null. Baseline levels of C-reactive protein (CRP) and the incidence and outcome of human sepsis. Nature Communications